Publications list

Below is a list of publications by our employees. Employees are listed alphabetically by family name. You can request copies by writing to info@natural-research.org

Kevin Duffy

Young, G.H. & Duffy, K. 1984.  ‘Night Herons in Scotland’.  Royal Zoological Society of Scotland Annual Report 1984. Edinburgh Zoo, Edinburgh.

Duffy, K. 1988. ‘Barn owls breeding near Edinburgh, 1985’.  Lothian Bird Report 1987.  Scottish Ornithologists Club, Lothian Branch.

Duffy, K. 1990.  ‘Breeding the Hawk-headed Parrot (Deroptyus acciptrinus acciptrinus) at Penscynor Wildlife Park’ Ratel, J. Association of British Wild Animal Keepers 17 (1): 14 –15.

Jones, C.G & Duffy, K. 1991. ‘Field work on the echo parakeet’. Psittascene, Newsl. World Parrot Trust 3 (4): 6 – 7.

Jones, C.G & Duffy, K. 1992. ‘Conservation of the echo parakeet Psittacula eques from Mauritius. Psittascene, Newsl. World Parrot Trust 4 (4): 7 - 10.

Jones, C.G & Duffy, K. 1993. ‘Conservation of the Echo Parakeet Psittacula eques echo’. J. Wildlife Preservation Trusts 29: 126 – 148.

Wotton, S.R., Carter, I., Cross, I.V., Etheridge, B.E. Snell, N.,  Duffy, K., Thorpe, R. & Gregory, R.D. 2002. ‘Breeding status of the Red Kite Milvus milvus in Britain in 2000. Bird Study 49, 278 – 286. British Trust for Ornithology, Thetford.

Evans, R., Broad, R., Duffy, K., MacLennan, A., Bainbridge, I. & Mudge, G. ‘Re-establishment of a population of White-tailed Eagles in Scotland. In: Helander, B., Marquiss, M. & Bowerman, W (eds.) 2003. SEA EAGLE 2000. Proceedings from an international conference at Bjorko, Sweden, 13 – 17 September 2000: 397 - 403. Swedish Soc. for Nature Conservation/SNF, Stockholm.

Bainbridge, I.P, Evans, R.J, Broad, R.A., Crooke, C.H., Duffy, K., Green, R.E. Love, J.A. & Mudge, G.P. Reintroduction of white-tailed eagles (Haliaeetus albicilla) to Scotland. In: Birds of Prey in a Changing Environment. Scottish Natural Heritage 2003: 393 - 406. Proceedings of an SNH/BTO conference held at the Scottish Police College, Tulliallen Castle, December 2000. Edinburgh.

Carter, I., Cross, A.V., Douse, A., Duffy, K., Etheridge, B.E., Grice, P.V., Newberry, P., Orr-Ewing, D.C., O’Toole, L., Simpson, D & Snell, N. ‘Reintroduction and conservation of the red kite (Milvus milvus) in Britain: current threats and prospects for future range expansion’. In: Birds of Prey in a Changing Environment. Scottish Natural Heritage 2003: 407 - 416. Proceedings of an SNH/BTO conference held at the Scottish Police College, Tulliallen Castle, December 2000. Edinburgh.

Digger Jackson

Jackson, D.B., 1994. Breeding dispersal and site-fidelity in three monogamous wader species in the Western Isles, UK. Ibis 136, 463-473.

Fuller, R. J. and Jackson, D. B., 1999. Changes in populations of breeding waders on the machair of North Uist, Scotland 1983 – 1998. Wader Study Group Bulletin 90, 47-55.

Jackson, D.B. and Green, R. E., 2000. The importance of the introduced hedgehog (Erinaceus europaeus) as a predator of the eggs of waders (Charadrii) on the machair in South Uist, Scotland. Biological Conservation 93, 33-348.

Jackson, D.B., 2001. Experimental removal of introduced hedgehogs improves wader nest success in the Western Isles, Scotland. Journal of Applied Ecology 38, 802-812.

O’Brien, M., Tharme, A. and Jackson, D., 2002. Changes in breeding wader numbers on Scottish farmed land during the 1990s. Scottish Birds 23, 10-21

Jackson D.B., 2003. Between-lake differences in the diet and provisioning behaviour of Black-throated Divers Gavia arctica breeding in Scotland. Ibis 145, 30-44.

Jackson D.B., 2003. Waders, hedgehogs and machair: research and conservation lessons from the Outer Hebrides. Wader Study Group Bulletin Vol. 100

Jackson, D.B., Fuller, R.J. and Campbell, S.T. 2004 Long-term population changes among breeding shorebirds in the Outer Hebrides, Scotland, in relation to an introduced mammalian predator. Biological Conservation 117, 151-166.

Jackson, D.B., 2005. Environmental correlates of lake occupancy and brood survival of Black-throated Divers Gavia arctica in Scotland. Bird Study 52, 225-236.

Jackson, D.B., 2006. The breeding biology of introduced hedgehogs on a Scottish island: lessons for population control and bird conservation. Journal of Zoology (London) 268, 303-314.

Jackson , D.B. In press. Factors affecting the abundance of introduced hedgehogs (Erinaceus europaeus) to the Hebridean island of South Uist in the absence of natural predators and the implications for nesting birds . Accepted Journal of Zoology (London) May 2006.

Jackson, D., and Jit, R. Submitted. Population density and detectability of three Fijian forest birds. Notornis May 2006.

Fiona Leckie

Leckie, F., Arroyo, B.E., Thirgood. S.J. & Redpath, S.M. (2008) Parental differences in brood provisioning by Hen Harriers Circus cyaneus. Bird Study.

Moss R, Piertney SB, McColl A, Lock K, Mougeot F, Leckie F, Redpath SM, Dallas J, Bacon P & Lambin X. (2007) The rise and fall of density, kinship and aggressiveness during a red grouse cycle. Ecology

Piertney SB, Lambin X, McColl A, Lock K, Bacon P, Dallas J, Mougeot F, Leckie F, Racey P, Redpath SM & Moss R. (2007) Temporal changes in kin structure and aggressiveness as a mechanism driving unstable population dynamics in red grouse (Lagopus lagopus scoticus). Ecology Letters

Mougeot F, Perez-Rodriguez L, Martinez-Padilla J & Redpath SM & Leckie F. (2007) Intestinal parasites and testosterone influence carotenoid signalling in a Tetraonid bird. J Evol Biol

Evans S, Mougeot F, Redpath SM & Leckie F. (2007) Alternative methods for estimating breeding density in an upland gamebird, the red grouse. Wildlife Biology.

Redpath, S.M., Mougeot, F., Leckie, F.M., Elston, D.A., and Hudson, P.J. (2006) Ecology Letters 9: 410-418.

Redpath S, Mougeot F, Leckie F & Evans S. (2006) The effects of autumn testosterone on survival and productivity in red grouse Lagopus lagopus scoticus. Animal Behaviour online.

Redpath, SM, Leckie, F., Arroyo, B., Amar, A. & Thirgood, S. (2006) Compensating for the costs of polygyny in hen harriers Circus cyaneus. Behavioural Ecology and Sociology (online)

Seivwright L, Redpath SM, Mougeot F, Leckie F & Hudson PJ . (2005) Interactions between intrinsic and extrinsic mechanisms in a cyclic species: testosterone increases parasite infection in red grouse. Proceedings of the Royal Society-Biological Sciences 172: 2299-2304.

Mougeot F, Redpath SM & Leckie F. (2005) Ultra-violet reflectance of male and female red grouse sexual ornaments reflects nematode parasite intensity. Journal of Avian Biology 36: 203-209.

Mougeot F, Piertney S, Leckie F, Evans S, Moss R, Redpath SM & Hudson PJ. (2005) Experimentally increased aggressiveness reduces population kin structure and subsequent recruitment in red grouse Lagopus lagopus scoticus. Journal of Animal Ecology 74: 488-497.

Mougeot F, Dawson A, Redpath SM & Leckie F. (2005) Testosterone and autumn territorial behaviour in male red grouse Lagopus lagopus scoticus. Hormones & Behavior 47: 576-584

Redpath, S.M., Arroyo, B.E., Leckie, F,M., Bacon, P., Bayfield, N., Gutiérrez, R.J. & Thirgood, S.J. (2004) Using decision modelling with stakeholders to reduce human-wildlife conflict: a raptor - grouse case study. Conservation Biology18: 350-359

Mougeot, F., Redpath, S.M., Leckie, F. and Hudson, P.J. (2003) The effect of aggressiveness on the population dynamics of a territorial bird. Nature 421, 737-739

Redpath, S.M., Arroyo, B.E., Etheridge, B., Leckie, F., Bouwman, K. and Thirgood, S.J. (2002) Temperature and hen harrier productivity: from local mechanisms to geographical patterns. Ecography 25, 533-540

Redpath, S. Amar, A., Madders, M., Leckie, F. and Thirgood, S. (2002) Hen harrier foraging success in relation to land use in Scotland. Animal Conservation 5, 113-118

Redpath, S.M., Thirgood, S.J. & Leckie, F. (2001) Does supplementary feeding reduce predation of Red Grouse by Hen Harriers? J. Appl. Ecol.38 (6)1157-1168

Leckie, F. and Campbell, S. (2000) A new record of successful tree nesting peregrines. Scottish Birds 21(1) 45-46

Leckie, F. M., Thirgood, S.J., May, R., & Redpath S. M. (1998) Variation in the diet of red foxes on Scottish moorland in relation to prey abundance. Ecography 21(6) 599-604

Madders, M., Leckie, F.M., Watson, J., & McKay, C.R. (1998) Distribution and foraging habitat preferences of Choughs on The Oa peninsula, Islay.  Scottish Birds 19 (5) 280-289.

Thirgood, S.J., Leckie, F.M. & Redpath, S.M.  (1996) Diurnal and seasonal variation in line transect counts of moorland passerines. Bird Study 42.

Mike McGrady

McGrady, M. J., Young, G. S. and Seegar, W. S. 2006. Migration of a Peregrine falcon Falco peregrinus over water in the vicinity of a hurricane. Ringing and Migration 23, 80 – 84.

Haworth, P. F., McGrady, M. J., Whitf.ield, D. P., Fielding, A. H. and McLeod, D. R. A. 2006. Ranging distance of adult Golden Eagles Aquila chrysaetos in western Scotland according to season and breeding status. Bird Study 53, 265-273.

McGrady, M. J. and Pennerstorfer, J. 2006. GIS-supported analyses of Golden Eagle (Aquila chrysaetos) habitats: Characteristics of some eagle ranges in Austria. Pp 61-68. In: Gamauf, A. and Berg, H.-M. (Eds.) Greifvögel and Eulen in Österreich. Naturhistorisches Museum, Wien, Vienna.

Gavashelishvili, A., M. J. McGrady and Z. Javakhishvili. 2006. Breeding site selection by bearded vulture (Gypaetus barbatus) and Eurasian griffon (Gyps fulvus) in the Caucasus. Animal Conservation 9: 159-170.

McGrady, M. J. and Gavashelishvili, A. 2006. Radiotracking vultures into Iran. Podoces 1 (1): 21-26.

Gavashelishvili, A. and M. J. McGrady. 2006. Some results of research on Cinereous vulture Aegypius monachus in Georgia. Vulture News 54:4-10.

Gavashelishvili, A., M. J. McGrady and Z. Javakhishvili. 2006. Breeding site selection by bearded vulture (Gypaetus barbatus) and Eurasian griffon (Gyps fulvus) in the Caucasus. Animal Conservation 9: 159-170.

Gavashelishvili, A. and M. J. McGrady. 2006. GIS-based modelling of vulture response to carcass appearance in the Caucasus. J. Zool. 269: 365-372.

Haworth, PF, McGrady, MJ, Whitfield, DP, Fielding, AH & McLeod, DRA. 2006. Ranging distance of adult golden eagles Aquila chrysaetos in western Scotland according to season and breeding status. Bird Study 53, 265-273

McGrady, M. J. and Pennerstorfer, J. In press. GIS-supported analyses of Golden Eagle (Aquila chrysaetos) habitats: Characteristics of some eagle ranges in Austria. Egretta (Accepted February 2005).

Gavashelishvili, A., McGrady, M. J. & Javakhishvili, Z. 2006. Planning the conservation of the breeding population of cinereous vultures (Aegypius monachus) in the Republic of Georgia. Oryx 40 (1): 1-8..

Peske, L. and McGrady, M. 2005. A system for locating satellite-received transmitters (PTTs) in the field. Wildl. Soc. Bull. 33(1): 307-312.

Walker, D., M. McGrady, A. McCluskie, M. Madders & D. R. A. McLeod. 2005. Resident Golden Eagle ranging behaviour before and after construction of a windfarm in Argyll. Scottish Birds 25 (1) 24-40.

McGrady, M. J. 2004. The golden eagle: the biology, ecology and management of a large avian predator. Keynote address. Proceedings of Symposium om kongeørn, Stjørdal, Nord-Trøndelag, Norway 3-5 October 2003.

Katzner, T., A. Gavashelishvili, S. Skylarenko, M. McGrady, J. Shergalin and K. Bildstein. 2004. Population and conservation status of griffon vultures in the former Soviet Union. Pp. 235-240 In: Meyburg, B.-U., R.D. Chancellor (Eds.) Raptors Worldwide. World Working Group on Birds of Prey, MME-Birdlife Hungary.

McGrady, M.J., Petty, S. J. and McLeod D.R.A. 2004. Potential impacts of new native woodland expansion on golden eagles in Scotland, Scottish Natural Heritage. Commissioned Report No. 018 (ROAME No. F99LD01).

Ganusevich, S. A., Maechtle, T. L., Seegar, W. S., Yates, M. A., McGrady, M. J., Fuller, M., Schueck, L., Dayton, J., Henny, C. J. 2004. Autumn migration and wintering areas of Peregrine Falcons (Falco peregrinus) nesting on the Kola Peninsula, northern Russia. Ibis 146: 291-297.

Haines, A. M., McGrady, M. J., Martell, M.S., Dayton, B. J., Henke, B., Seegar, W.S. 2003. Migration routes and wintering locations of Broad-winged Hawks tracked via satellite Wilson Bull. 115: 166-169.

McGrady, M. J., S. J. Petty, I. P. Bainbridge, D. R. A. McLeod. 2003. Golden eagles and new native woodlands: the role of the Woodland Grant Scheme in Scotland. Pages 341-350. In: Thompson D. B. A. (Ed.) Birds of Prey in a Changing Environment. The Stationary Office, Edinburgh.

Fielding, A., D. McLeod, M. McGrady, P. Whitfield and P. Haworth. 2003. Modelling the impact of land use change on golden eagles Aquila chrysaetos Pages 323-340. In: Thompson D. B. A. (Ed.) Birds of Prey in a Changing Environment. The Stationary Office, Edinburgh.

Ueta, M., McGrady, M.J. Nakagawa, H., Sato, F., and Masterov, V. 2003. Seasonal change in habitat use in Steller’s sea eagles. Oryx 37(1): 110-114.

McGrady, M. J., M. Ueta, E. Potapov, I. Utekhina, V. Masterov, A. Ladyguine, V. Zykov, J. Cibor, M. Fuller, W.S. Seegar. 2003. Migration and wintering of juvenile and immature Steller’s sea eagles. Ibis 145: 318-328.

McLeod, D. R. A., Whitfield, D. P., Fielding, A. H., Haworth, P., McGrady, M. J. 2003. Predicting home range use by golden eagles Aquila chrysaetos in western Scotland. Avian Science 2: 183-198.

McGrady, M. J., T. L. Maechtle, J. J. Vargas, W. S. Seegar, C. Porras Peña, 2002. Movements of peregrine falcons Falco peregrinus wintering on the Gulf Coast of Mexico 1996-1998. Condor 104: 39-48.

McGrady, M. J., J. R. Grant, I. P. Bainbridge, D. R. A. McLeod. 2002. A model of golden eagle (Aquila chrysaetos) ranging behavior. Pp. 62-69. In M.J. McGrady, R. Rodríquez-Estrella, M.J. Bechard, and E. Iñigo-Elias (Eds.) The golden eagle: Its status, conservation and management in 1999. Journal of Raptor Research 36 (1 Suppl.), March 2002.

McLeod, D.R.A., D.P. Whitfield, M.J. McGrady. 2002. Improving prediction of golden eagle (Aquila chrysaetos) ranging in western Scotland using GIS and terrain modelling. Pp 70-77. In M.J. McGrady, R. Rodríquez-Estrella, M.J. Bechard, and E. Iñigo-Elias (Eds.) The golden eagle: Its status, conservation and management in 1999. Journal of Raptor Research 36 (1 Suppl.), March 2002

McGrady, M. J., R. Rodríquez-Estrella, M.J. Bechard, and E. Iñigo-Elias (Eds.) 2002. The golden eagle: Its status, conservation and management in 1999. Journal of Raptor Research 36 (1 Suppl.).

Gavashelishvili, A. and McGrady, M. J. 2002. Conservation of Gyps vultures in Asia: reports from the workshop. 3rd North American Ornithological Conference. 24-28 September 2002. New Orleans, Louisiana, USA.

Gavashelishvili, A. and M. J. McGrady. 2002. Preliminary results of a predictive model for breeding site selection by the bearded vulture (Gypaetus barbatus) in the Caucasus. Pages 87-94. In: Sakoulis, A., M. Probonas & S. Xirouchakis (Eds.) Proceedings of the 4th Bearded Vulture Workshop, Natural History Museum of Crete, University of Crete, Greece.

Gavashelishvili, A. and M. J. McGrady. 2002. Vultures in Georgia. Pp 18-19. In: T. Katzner and J. Parry-Jones (Eds.). Conservation of Gyps vultures in Asia. Workshop proceedings. 3rd North American Ornithological Conference 24-28 September 2002 New Orleans, Louisiana, USA.

McGrady, M. J., Petty, S. J., Mudge, G and Bainbridge, I. P. 2001. Guidance for the establishment and management of new native woodlands in Scotland to promote golden eagle (Aquila chrysaetos) conservation. Abstracts of 4th Eurasian Congress on Raptors. Seville 25-29 September 2001. p. 123.

Potapov, E., McGrady, M., Utekhina, I., and Garcelon, D. 2001. Numbers and trends in Steller’s sea eagle: a review. Abstracts of 4th Eurasian Congress on Raptors. Seville 25-29 September 2001. p. 147.

McGrady, M. J., M. J. Ueta, E. R. Potopov, I. Utekhina, V. B. Masterov, M. Fuller, W. S. Seegar, A. Ladyguin, E. G. Lobkov & V. B. Zykov. 2000. Migration and wintering of juvenile and immature Steller’s sea eagles. Pp. 83-90. In: Ueta, M. J. and M. J. McGrady (Eds.) First Symposium on Steller’s and White-tailed Sea Eagles in East Asia. Wild Bird Society of Japan. Tokyo.

Potopov, E. R., I. Utekhina & M. J. McGrady 2000. Steller’s sea eagle in Magadan District and in the north of Khabarovsk District. Pp. 29-44. In: Ueta, M. J. and M. J. McGrady (Eds.) First Symposium on Steller’s and White-tailed Sea Eagles in East Asia. Wild Bird Society of Japan. Tokyo.

Utekhina, I., E. Potopov & M. J. McGrady 2000. Diet of the Steller’s sea eagle in the northern Sea of Okhotsk. Pp 71-82. In: Ueta, M. J. and M. J. McGrady (Eds.) First Symposium on Steller’s and White-tailed Sea Eagles in East Asia. Wild Bird Society of Japan. Tokyo.

McGrady, M., Potapov, E., and Utekhina, I. 1999. Feeding on Steller’s sea eagle (Haliaeetus pelagicus) nestling by a brown bear (Ursus arctos). Journal of Raptor Research 33 (4): 342-343.

Grant J. R. and McGrady, M. J. 1999. Dispersal of golden eagles in Scotland. Ringing and Migration 19: 160-174.

McGrady, M.J. 1998. Models of golden eagle home ranges, a response to Walker. Scottish Bird News 50: 5-6.

McGrady, M.J. 1998. The Golden Eagle by Jeff Watson, an extended review. Scottish Bird News 50: 4.

McGrady, M. J., J. R. Grant, D.R.A. McLeod. 1998. A preliminary analysis of the relation between land cover and golden eagle Aquila chrysaetos ranging behaviour in Argyll western Scotland. Pages 263-270. In: Meyburg, B.-U., R.D. Chancellor, and J.J. Ferrero (Eds.) Holarctic Birds of Prey, WWGBP and ADNEX. Berlin.

McGrady, M. J. 1998. Radio-tracking of Steller’s Sea Eagle Yields Surprises. Russian Conservation News 14.

McGrady, M. J., McLeod, D. M., S.M Petty, J. R. Grant and I. P. Bainbridge. 1997. Eagles and forestry. Forestry Commission Research Information Note. No. 292. HMSO. London.

McGrady, M.J. 1997. Golden eagle. BWP Update 1(2): 99-114.

McGrady, M. J. and J. R. Grant. 1996. A modified 'power snare' for the capture of breeding golden eagles (Aquila chrysaetos). Journal of Raptor Research 30(1): 28-31.

Peske, L. and McGrady, M. J. 1995. Unusual prey for golden eagle Aquila chrysaetos. Buteo 7: 75-77. (In Czech).

McGrady, M. J. D. C. Orr-Ewing, and T. J. Stowe 1994. The re-introduction of the red kite Milvus milvus into Scotland. Pages 471-477. In: Meyburg, B. -U. and R. D. Chancellor. Editors. Raptor Conservation Today. World Working Group on Birds of Prey and Pica Press.

Green, M., M. McGrady, S. Newton, and J. Uttley. 1994. Counts of shorebirds of Barr al Hikman and Ghubbat al Hashish, Oman, winter 1989-90. Wader Study Group Bulletin 72: 39-43.

Clark, W., K. Duffy, E. Gorney, M. McGrady, and C. Schultz. 1986. Raptor ringing in Eilat, Israel. Sandgrouse 7: 21-28.

Ruth Tingay

Tingay, R.E., Culver, M. and Watson, R.T. (In press). Using molecular sexing to assess field-based techniques for gender assignment in the Madagascar fish eagle Haliaeetus vociferoides. Journal of Raptor Research.

Tingay, R.E., Dawson, D.A., Pandhal, J., Clarke, M.L., David, V., Hailer, F. and Culver, M. (2007). Isolation of 22 new Haliaeetus microsatellite loci and their characterisation in the critically endangered Madagascar fish eagle Haliaeetus vociferoides and three other Haliaeetus eagle species. Molecular Ecology Notes: on-line early [January 2007].

Tingay, R.E., Dawson, D.A., Hailer, F., Pandhal, J., Clarke, M.L. and Culver, M. (submitted). Low genetic variability in the critically endangered Madagascar fish eagle Haliaeetus vociferoides in contrast to three other Haliaeetus eagle species.

Tingay, R.E., Nicoll, M.A.C. and Sun, V. (2006). Status and distribution of the grey-headed fishing eagle Ichthyophaga ichthyaetus in the Prek Toal Area of Tonle Sap Lake, Cambodia. Journal of Raptor Research 40 (4): 277-283.

Tingay, R.E., Culver, M. and Watson, R.T. (2006). Conservation genetics of the Madagascar fish eagle (Haliaeetus vociferoides). Abstracts, American Ornithologists Union Conference, October 2006, Veracruz, Mexico.

Tingay, R.E. (2005). Historical distribution, contemporary status and cooperative breeding in the Madagascar fish eagle Haliaeetus vociferoides: implications for conservation. PhD dissertation, University of Nottingham, UK. Pp. 413.

Tingay, R.E. and Clarke, M. (2004). Politics at the nest: unusual reproductive strategies amongst cooperatively breeding Madagascar fish eagles Haliaeetus vociferoides. Abstracts, Raptor Research Foundation Annual Conference, Bakersfield, California.

Tingay, R.E., Clarke, M. and Culver, M. (2004). Re-evaluating the conservation status of the critically endangered Madagascar fish eagle Haliaeetus vociferoides: historical population decline or a naturally small population? Abstracts, Raptor Research Foundation Annual Conference, November 2004, Bakersfield, California.

Tingay, R.E., Clarke, M., Watson, R.T., Thorstrom, R. and Kalavaha, L. (2004). Survival and behaviour of a one-footed Madagascar fish eagle in the wild. Journal of Raptor Research 38: 85-88.

Tingay, R.E. (2002). Three years with Madagascar fish eagles. The Peregrine Fund Newsletter 33: 14-16.

Tingay, R.E., Culver, M., Hallerman, E., Watson, R.T. and Fraser, J.D. (2002). Subordinate Madagascar fish eagle Haliaeetus vociferoides males sire young in polyandrous breeding trios. Journal of Raptor Research 36: 280-286.

Gilbert, M. and Tingay, R.E. (2001). Predation of a fat-tailed dwarf lemur Cheirogaleus medius by a Madagascar harrier-hawk Polyboroides radiatus: an incidental observation. IUCN Primate Specialist Group Newsletter 6: 6.

Tingay, R.E., Hallerman, E., Culver, M. and Watson, R.T. (2000). Sex, lies and dominance: behaviour and paternity of extra-pair Madagascar fish eagles Haliaeetus vociferoides. Abstracts, Raptor Research Foundation Annual Conference, Jonesboro, Arkansas.

Tingay, R.E. and Gilbert, M. (2000). Behaviour of banded kestrel Falco zoniventris in western Madagascar: a possible foraging association with sickle-billed vanga Falculea palliata. Bulletin of the African Bird Club 7: 111-113.

Ruelas, E., Hoffman, S.W., Goodrich, L.J. and Tingay, R.E. (2000). Conservation strategies for the world’s largest known raptor migration flyway: Veracruz, The River of Raptors. In Chancellor, R.D. and Meyburg, B.-U. (Eds.). Raptors At Risk: Proceedings of the V World Conference on Birds of Prey and Owls, August 1998, Johannesburg, South Africa. WWGBP/Hancock House, Surrey, UK. Pp. 591-596.

Tingay, R.E. (2000). Sex, lies and dominance: behaviour and paternity of extra-pair Madagascar fish eagles. MSc dissertation, University of Nottingham, UK.

Tingay, R.E. and Gilbert, M. (1999). Annotated list of the birds of Lacs Soamalipo, Befotaka and Ankerika and surrounding Tsimembo Forest, western Madagascar. Working Group on Birds in the Madagascar Region Newsletter 9: 26-30.

Tingay, R.E. (1998). Veracruz: A River of Raptors. Hawk & Owl Trust Annual Review 25: 17-21.

Tingay, R.E., Atkinson, E.C. and Bildstein, K.L. (1996). Philopatry and inter-year nest site fidelity in American kestrels Falco sparverius. Abstracts, Raptor Research Foundation Annual Conference, Boise, Idaho.

Dave Walker

Walker, D.G. 1987 Observations on the post-fledging period of the Golden Eagle Aquila chrysaetos in England. Ibis 129:92-96

Walker, D.G. 1988 The behaviour and movements of a juvenile Golden Eagle Aquila chrysaetos in England. Ibis 130:564-565

Walker D.G.1983 Golden Eagle killing mobbing Carrion Crows. Brit. Birds 76:309-310

Walker D.G. 1988 Unusual Golden Eagle Call. Scott. Bird News 9.8.

Holmes J., Walker D., Davies P. and Carter I. 2000. The illegal persecution of raptors in England. 2000. EN Research Report No 343

Walker, D.G. 2000. Food Supply and Breeding Performance of the Haweswater Golden Eagles. 2001. Birds and Wildlife in Cumbria: 65-68. Cumb. Nats. Union

Walker, D.G. 2003. Some notes on the Red Deer (Cervus elaphus) at Haweswater, Mardale. The Carlisle Naturalist: 11:No 1: 14-16

Walker, D.G. 2004 The Lakeland Eagles. ISBN 0 9517378 0 5

Walker, D., M. McGrady, A. McCluskie, M. Madders & D. R. A. McLeod. 2005. Resident Golden Eagle ranging behaviour before and after construction of a windfarm in Argyll. Scottish Birds 25 (1) 24-40.

Phil Whitfield

Whitfield, DP. 1985. Raptor predation on wintering waders in southeast Scotland. Ibis 127: 544-558.

Whitfield, DP. 1986. Plumage variability and territoriality in breeding turnstone Arenaria interpres: status signalling or individual recognition? Animal Behaviour 34: 1471-1482.

Whitfield, DP. 1987. Plumage variability, status signalling and individual recognition in avian flocks. Trends in Ecology and Evolution 2: 13-17.

Whitfield, DP & Magnusson, J. 1987. The migration of high arctic waders through Melrakkasletta, northeast Iceland. Wader Study Group Bulletin, 49 Suppl./IWRB Special Publication 7: 85-89.

Whitfield, DP. 1988. The social significance of plumage variability in wintering turnstone Arenaria interpres. Animal Behaviour 36: 408-415.

Whitfield, DP. 1988. Sparrowhawks Accipiter nisus affect the spacing behaviour of wintering turnstone Arenaria interpres and redshank Tringa totanus. Ibis 130: 284-287.

Summers, RW, Corse, CJ, Nicoll, M, Smith, R & Whitfield, DP. 1988. The biometrics and wintering area of Icelandic purple sandpipers. Ringing & Migration 9: 133-138.

Whitfield, DP, Evans AD & Whitfield PA. 1988. The impact of raptor predation on wintering waders. Proceedings of the International Ornithological Congress 19: 674-687.

Whitfield, DP. 1990. Individual feeding specialisations in wintering turnstone Arenaria interpres. Journal of Animal Ecology 59: 193-211.

Whitfield, DP. 1990. Male choice and sperm competition as constraints on polyandry in the red-necked phalarope Phalaropus lobatus. Behavioural Ecology & Sociobiology 27: 247-254.

Whitfield, DP & Brade, JJ. 1991. The breeding behaviour of the Knot Calidris canutus. Ibis 133: 246-255.

Galbraith, H, Murray, S, Duncan K, Smith, R, Whitfield, DP & Thompson DBA. 1993. Diet and habitat use of the Dotterel Charadrius morinellus in Scotland. Ibis 135: 148-155.

Galbraith, H, Murray, S, Rae, S, Whitfield, DP & Thompson, DBA. 1993. Numbers and distribution of the dotterel (Charadrius morinellus) breeding in Great Britain. Bird Study: 40: 161-169.

Cresswell, W & Whitfield, DP. 1994. The effects of raptor predation on wintering wader populations at the Tyninghame estuary, southeast Scotland. Ibis 136: 223-232.

Whitfield, DP. 1995. Behaviour and ecology of a polyandrous population of Grey Phalaropes Phalaropus fulicarius in Iceland. Journal of Avian Biology 26: 349-352.

Smith, R & Whitfield, DP. 1995. Renesting by male Dotterel Charadrius morinellus after successfully rearing chicks. Bird Study 42: 174-175.

Whitfield, DP & Tomkovich, PS. 1996. Mating system and timing of breeding in Holarctic waders. Biological Journal of the Linnaean Society 57: 277-290.

Whitfield, DP, Brade, JJ, Burton, RW, Hankinson, KW & Young, S. 1996. The abundance of breeding Knot Calidris canutus islandica. Bird Study 43: 290-299.

Whitfield, DP, Duncan K, Pullan, D & Smith, RD. 1996. Recoveries of Scottish-ringed Dotterel Charadrius morinellus in the non-breeding season: evidence for seasonal shifts in wintering distribution. Ringing & Migration 17: 105-110.

Whitfield, DP. 1997. Waders (Charadrii) on Scotland’s blanket bogs: recent changes in numbers of breeding birds. In: Conserving Peatlands (Eds. L Parkyn, RE Stoneman & HAP Ingram), pp. 103-111. CAP International, Wallingford.

Downie, IS, Foster, GN, Coulson, JC & Whitfield, DP. 1998. The water beetles of Badanloch Bog, Sutherland. Glasgow Naturalist 23: 31-34.

Downie, IS, Foster, GN, Coulson, JC & Whitfield, DP. 1998. Distribution of aquatic macroinvertebrates within peatland pool complexes in the Flow Country, Scotland. Hydrobiologia. 377: 95-105.

Whitfield, DP. 1999. Methods for ageing Dotterel Charadrius morinellus. Ringing & Migration 19: 200-204.

Whitfield, DP, Cresswell, W, Ashmole, NP, Clark, NA & Evans, AD. 1999. No evidence for sparrowhawks selecting redshanks according to size or condition. Journal of Avian Biology 30: 31-39.

Fuller, RJ, Gillings, S & Whitfield, DP. 1999. Responses of breeding birds to expansion of scrub in the eastern Scottish Highlands: preliminary implications for conservation strategies. Vogelwelt 120 Suppl.: 53-62.

Whitfield, DP, McLeod, DRA, Fielding, AH, Evans, RJ, Broad, RA & Haworth, PF. 2001. The effects of forestry on golden eagles on the island of Mull, western Scotland. Journal of Applied Ecology 38: 1208-1220.

Whitfield, DP. 2002. Status of breeding Dotterel Charadrius morinellus in Britain in 1999. Bird Study 49: 237-249.

Whitfield, DP, Evans, RJ, Broad, RA, Fielding, AH, Haworth, PF, Madders, M & McLeod, DRA. 2002. Are reintroduced white-tailed eagles in competition with golden eagles? Scottish Birds 23: 36-45.

McLeod, D, Whitfield, DP & McGrady, MJ. 2002. Improving prediction of golden eagle (Aquila chrysaetos) ranging in western Scotland, using GIS and terrain modelling. Journal of Raptor Research 36 (1 Supplement): 70-77.

Watson J & Whitfield, P. 2002. A conservation framework for the golden eagle Aquila chrysaetos in Scotland. Journal of Raptor Research 36 (1 Supplement): 41-49.

Holt, S, Whitfield, DP & Gordon, J. 2002. Potential reproductive rates in the Eurasian dotterel. Bird Study 49: 87-88.

Reid, JM, Cresswell, W, Holt, S, Mellanby, RJ, Whitfield, DP & Ruxton, GD. 2002. Nest scrape design and clutch heat loss in Pectoral Sandpipers Calidris melanotos. Functional Ecology 16: 305-312.

Holt, S, Whitfield, DP, Duncan, K, Rae, S & Smith, RD. 2002. Mass loss in incubating Eurasian dotterel: adaptation or constraint? Journal of Avian Biology 33:219-224.

McLeod, DRA, Whitfield, DP, Fielding, AH, Haworth, PF & McGrady, MJ. 2002. Predicting home range use by golden eagles Aquila chrysaetos in western Scotland. Avian Science 2: 183-198.

Buchanan, GM, Pearce-Higgins, JW, Grant, MC & Whitfield, DP. 2003. Correlates of the change in Ring Ouzel Turdus torquatus abundance in Scotland from 1989-91 to 1999. Bird Study 50: 97-105.

Whitfield, DP. 2003. Predation by Eurasian sparrowhawks produces density-dependent mortality in wintering redshanks. Journal of Animal Ecology 72: 27-35.

Whitfield, DP, McLeod, DRA, Watson, J, Fielding, AH & Haworth, PF. 2003. The association of grouse moors in Scotland with the illegal use of poisons to control predators. Biological Conservation 114: 157-163.

Cresswell, W, Holt, S, Reid, JM, Whitfield, DP & Mellanby, RJ. 2003. Do energetic demands constrain incubation scheduling in a biparental species? Behavioral Ecology 14: 97-102.

Holt, S & Whitfield, DP. 2003. Timing of nest departure by broods of dotterel Charadrius morinellus. Bird Study 49: 88-90.

Whitfield, DP. 2003. Redshank Tringa totanus flocking behaviour, distance from cover and vulnerability to sparrowhawk Accipiter nisus predation. Journal of Avian Biology 34: 163-169.

Whitfield, DP. 2003. Density-dependent mortality of wintering Dunlins Calidris alpina through predation by Eurasian Sparrowhawks Accipiter nisus. Ibis 145: 432-438.

Whitfield, DP. 2003. Raptor predation on non-breeding shorebirds: some thoughts for the future. Wader Study Group Bulletin 100: 134-137.

Thompson, DBA, Whitfield, DP, Galbraith, H, Duncan, K, Smith, RD, Murray, S & Holt, S. 2003. Breeding bird assemblages of alpine areas in Scotland. In: Nagy, L, Grubher, G, Korner, C & Thompson, DBA (eds.) Alpine Biodiversity in Europe. Springer-Verlag, London.

Whitfield, DP. 2003. Does timing of breeding affect breeding plumage acquisition in shorebirds? Wader Study Group Bulletin 101/102: 77-80.

Whitfield, DP. 2003. The effects of predation by Sparrowhawks on the behaviour and population dynamics of wintering Redshanks. In: Thompson, DBA, Redpath, SM, Fielding, AH, Marquiss, M & Galbraith, CA (eds.), pp. 269-274. Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh.

Watson, J, Fielding, AH, Whitfield, DP, Broad, RA, Haworth, P, Nellist, K, Crane, K & Macdonald, EJ. 2003. Golden eagle breeding performance in relation to climate in western Scotland during the period 1981-2000. In: Thompson, DBA, Redpath, SM, Fielding, AH, Marquiss, M & Galbraith, CA (eds.), pp. 149-161. Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh.

Fielding, A, Whitfield, DP, McLeod, DRA, McGrady, MJ, & Haworth, PF. 2003. Modelling the impact of land use change on golden eagles (Aquila chrysaetos). In: Thompson, DBA, Redpath, SM, Fielding, AH, Marquiss, M & Galbraith, CA (eds.), pp. 323-340. Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh.

Fielding, AH, Haworth, PF, Morgan, DH, Thompson, DBA & Whitfield, DP. 2003. The impact of golden eagles (Aquila chrysaetos) on a diverse bird of prey assemblage. In: Thompson, DBA, Redpath, SM, Fielding, AH, Marquiss, M & Galbraith, CA (eds.), pp. 221-244. Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh.

Cresswell, W, Holt, S, Reid, JM, Whitfield, DP, Mellanby, RJ, Norton, D & Waldron, S. 2004. The energetic costs of egg heating constrain incubation attendance but do not determine daily energy expenditure in the pectoral sandpiper. Behavioral Ecology 15: 498-507.

Mee, A, Whitfield, DP, Thompson, DBA & Burke, TA. 2004. Extra-pair paternity in the common sandpiper Actitis hypoleucos, revealed by DNA fingerprinting. Animal Behaviour 67, 333-342.

Whitfield, DP, Fielding, AH, McLeod, DRA & Haworth, PF. 2004. The effects of persecution on age of breeding and territory occupation in golden eagles in Scotland. Biological Conservation 118: 249-259.

Whitfield, DP, Fielding, AH, McLeod, DRA & Haworth, PF. 2004. Modelling the effects of persecution on the population dynamics of golden eagles in Scotland. Biological Conservation 119, 319-333.

Whitfield, DP. In press. Territory holders and satellites in wintering redshanks Tringa totanus: ageism and queue jumping. Ibis.

Haworth, PF, McGrady, MJ, Whitfield, DP, Fielding, AH & McLeod, DRA. 2006. Ranging distance of adult golden eagles Aquila chrysaetos in western Scotland according to season and breeding status. Bird Study 53, 265-273

Madders, M & Whitfield, DP. 2006. Upland raptors and the assessment of windfarm impacts. Ibis 148 Supplement, 43-56.

Whitfield, DP, Fielding, AH, Gregory, MJP, Gordon AG, McLeod, DRA & Haworth, PF. 2007. Complex effects of habitat loss in the golden eagle Aquila chrysaetos. Ibis 149, 26-36.

Whitfield, DP. Individual differences in susceptibility to foraging competition and predation in wintering redshanks. (Under review for Journal of Animal Ecology.)

Eaton, M.A., Dillon, I.A., Stirling-Aird, P.K. & Whitfield, DP. In press. The status of the Golden Eagle Aquila chrysaetos in Britain in 2003. Bird Study.

Madders, M., Whitfield, DP & McCluskie, A. In press. The impacts of wind farms on hen harriers Circus cyaneus: implications for assessment. In: Proceedings of BES-IEEM Conference, EcIA: Science and Best Practice, University of Bath, July, 2006.

Fielding, AH, Whitfield, DP & McLeod, DRA. 2006. Spatial association as an indicator of the potential for future interactions between wind energy developments and golden eagles Aquila chrysaetos in Scotland. Biological Conservation 131, 359-369.

Whitfield, DP, Fielding, AH, McLeod, DRA, Morton, K, Stirling-Aird, P & Eaton, M. Using temporal change to identify spatial constraint influence on Golden Eagles Aquila chrysaetos in Scotland.

Whitfield, DP, Fielding, AH, McLeod, DRA, Haworth, PF & Watson, J. 2006. A conservation framework for the golden eagle in Scotland: refining condition targets and assessment of constraint influences. Biological Conservation 130, 465-480.

Whitfield, DP, MacColl, ADC, Holt, S, Denny, R, Gordon, J, Pullan, D, & Smith, R. The influence of human recreation on Eurasian Dotterel Charadrius morinellus breeding in the Scottish Highlands. I. The effect of disturbance. (Under review for Journal of Applied Ecology.)

Holt, S, Whitfield, DP, Denny, R, Dillon, I, Duncan, K, Gordon, J, Mee, A, Pullan, D, Rae, S, Smith, RD & Stirling, J. Energetic constraints and reproductive trade-offs in predator responses of incubating Eurasian Dotterel Charadrius morinellus (Under review for Animal Behaviour.)

Holt, S, Whitfield, DP & Bryant, D. Flexible incubation in the Eurasian Dotterel (Under review for Ibis.)

Holt, S, Whitfield, DP & Bryant, D. Does fine-scale nest site selection in the Eurasian dotterel minimize energetic costs or predation risk? (Under review for Behavioral Ecology.)

Cresswell, W, Lind, J, Quinn, JL, Minderman, J & Whitfield, DP. In press. Ringing or colour-banding does not increase predation risk. Journal of Avian Biology.

MacColl, ADC, Whitfield, DP, Denny, R, Gordon, J & Greene, D. Some patterns in the behaviour of recreational users of the Cairn Gorm plateau and other mountain areas in Scotland. (Under review for Scottish Geographical Magazine.)

MacColl, ADC, Whitfield, DP, Holt, S, Denny, R, Gordon, J, Pullan, D, Smith, R & Greene, D. Temporal trends in the recreational use of the Cairn Gorm plateau, Scotland. (Under review for Scottish Geographical Magazine.)

Whitfield, DP, Duncan, K & Holt, S. The influence of human recreation on Eurasian Dotterel Charadrius morinellus breeding in the Scottish Highlands. II. The attraction of predators. (First draft completed.)

Whitfield, DP & Partridge, L. Individual differences in the feeding behaviour of turnstone Arenaria interpres. I. Causes. (First draft completed.)

Whitfield, DP & Partridge, L. Individual differences in the feeding behaviour of turnstone Arenaria interpres. II. Consequences. (First draft completed.)

Whitfield, DP. 1989. Sexual conflict in the red-necked phalarope. Bliki 7: 59-62.

Whitfield, DP. 1996. Habitat requirements of breeding waders on blanket bogs in Northern Scotland. SNH Information and Advisory Note 53, Battleby.

Whitfield, DP. 1998. Environmental impacts of mountaineering and hillwalking: montane birds. SNH, Edinburgh.

Whitfield, DP. 1998. Red deer and sheep in the montane zone of Scotland. SNH, Edinburgh.

Whitfield, DP. 2000. Golden eagle Aquila chrysaetos ecology and conservation issues. SNH Review 132, Battleby

Whitfield, DP. 2002. Dotterel Charadrius morinellus. In The Migration Atlas: Movements of Birds of Britain and Ireland (eds C.V. Wernham, M.P. Toms, J. Marchant, J.A. Clark, G.M. Siriwardena & S.R. Baillie). Poyser, London.

Whitfield, DP. 2002. Turnstone Arenaria interpres. In The Migration Atlas: Movements of Birds of Britain and Ireland (eds C.V. Wernham, M.P. Toms, J. Marchant, J.A. Clark, G.M. Siriwardena & S.R. Baillie). Poyser, London.

Whitfield, DP & Brade, JJ. 1990. The waders of Borup Fiord. In: Report of the Joint Services Expedition to Borup Fiord, Ellesmere Island, 1988 (Ed. KW Hankinson). JSE Borup Fiord, London.

Whitfield, DP & Tharme, AP. 1997. An evaluation of the effects of native pinewood regeneration on the birds of the Cairngorms. SNH, Edinburgh.

Whitfield, DP, Evans, AD & Magnusson, J. 1989. The first confirmed breeding attempt of the Curlew Numenius arquata in Iceland. Bliki 8: 3-6.

Whitfield, DP, Duncan, K, Murray, S, Rae, S, Smith, R & Thompson, DBA. 1991. Monitoring the dotterel population of Britain. In: Britain's Birds in 1989-1990: the conservation and monitoring review (Eds. D Stroud & D Glue), pp 109-11. NCC, Peterborough.

Holt, S & Whitfield, DP. 1996. Montane Ecology Project site report series: Glas Maol, eastern Grampians. SNH, Edinburgh.

Holt, S & Whitfield, DP. 1996. Montane Ecology Project site report series: East Drumochter, central Grampians. SNH, Edinburgh.

Holt, S & Whitfield, DP. 1996. Montane Ecology Project site report series: Cairn Gorm study area. SNH, Edinburgh.

Tharme, AP & Whitfield, DP. 1996. Ornithological importance of moorland within the Forest of Strathspey and Forest of Mar Priority Forest Areas. RSPB, Edinburgh.

Thompson, DBA & Whitfield, DP. 1993. Research Progress Report: Research on mountain birds and their habitats. Scottish Birds 17: 1-8.

Thompson, DBA & Whitfield, DP. 1994. The Dotterel. In: Gibbons, DW, Reid, J & Chapman, R, The New Atlas of Breeding Birds in Britain and Ireland. Poyser, London.

McLeod, D. & Whitfield, DP 1999. GIS for Golden Eagle Management. SNH, Edinburgh.

Summers, RW, Corse CJ & Whitfield, DP. 1987. Purple Sandpiper studies in north Iceland 1986. Orkney Ringing Group, Kirkwall.

Whitfield, DP, Fielding, AH, Gregory, MJP, Gordon AG, McLeod, DRA & Haworth, PF. 2007. Complex effects of habitat loss on golden eagles Aquila chrysaetos. Ibis 149, 26-36.

Band, W, Madders, M, & Whitfield, DP. 2007. Developing field and analytical methods to assess avian collision risk at wind farms. In: de Lucas, M, Janss, GFE & Ferrer, M (eds.) Birds and Wind Farms: Risk Assessment and Mitigation, pp. 259-275. Quercus, Madrid.

Whitfield, DP, Fielding, AH, McLeod, DRA, Morton, K, Stirling-Aird, P & Eaton, M. 2007. Factors constraining the distribution of Golden Eagles Aquila chrysaetos in Scotland. Bird Study 54, 199-211.

Eaton, MA, Dillon, IA, Stirling-Aird, PK. & Whitfield, DP. 2007. Status of Golden Eagle Aquila chrysaetos in Britain in 2003. Bird Study 54, 212-220.

Downie, IS, Coulson, JC, O’Connell, MJ, Thomas, CJ, Evans, PR & Whitfield, DP. In press. Functional ecology of peatland animals in the Flow Country of northern Scotland: II. Invertebrate distribution and abundance. SNH Research, Survey & Monitoring Series, Battleby.

O’Connell, MJ, Thomas, CJ, Twiss, S, Downie, IS, Coulson, JC , Evans, PR & Whitfield, DP. In press. Functional ecology of peatland animals in the Flow Country of northern Scotland: I. Habitat use of breeding waders. SNH Research, Survey & Monitoring Series, Battleby.

O’Connell, MJ, Thomas, CJ, & Whitfield, DP. In press. Assessment of the abundance of birds in The Peatlands SPA using field survey and GIS-generated estimates. SNH Research, Survey & Monitoring Series, Battleby.

Parrott, D., Henderson, I., Deppe, C. & Whitfield, P. In press. Scottish racing pigeons killed by Peregrine Falcons Falco peregrinus: estimation of numbers from ring recoveries and Peregrine daily food intake. Bird Study

Whitfield, DP & Coupar, A. In press. Effects of terrestrial wind farms on birds and habitats. In: Baxter, J. & Band, W. (eds). Energy and the Natural Heritage. The Stationery Office, Edinburgh.

The above listing for Phil does not include 18 SNH reports on survey of upland and peatland birds in Lewis, Sutherland, Caithness, Shetland, the Cairngorms and several montane plateaux in Scotland.