Publication List

Below is a list of peer-reviewed scientific publications by our employees, listed in alphabetical order. Due to journal publishing regulations, we are not permitted to include PDF copies of our publications here. However, you can request PDF copies (for private use) by emailing us at:  In most cases the copyright for publications will be held by the author(s), not NR/NRP.  We will forward your request to the authors who will have discretion over whether to provide a copy of the publication. 

Bainbridge, I.P., Evans, R.J., Broad, R.A., Crooke, C.H., Duffy, K., Green, R.E., Love, J.A. & Mudge, G.P. 2003. Reintroduction of white-tailed eagles (Haliaeetus albicilla) to Scotland. In Thompson, D.B.A., Redpath, S.M., Fielding, A.H., Marquiss, M. & Galbraith, C.A. (Eds.). Birds of Prey in a Changing Environment. The Stationary Office, Edinburgh. Pp. 393-406.

Band, W., Madders, M. & Whitfield, D.P. 2007. Developing field and analytical methods to assess avian collision risk at wind farms. In de Lucas, M., Janss, G.F.E. & Ferrer, M. (Eds.). Birds and Wind Farms: Risk Assessment and Mitigation. Quercus, Madrid. Pp. 259-275.

Buchanan, G.M., Pearce-Higgins, J.W., Grant, M.C. & Whitfield, D.P. 2003. Correlates of the change in ring ouzel Turdus torquatus abundance in Scotland from 1989-91 to 1999. Bird Study 50: 97-105.

Carter, I., Cross, A.V., Douse, A., Duffy, K., Etheridge, B.E., Grice, P.V., Newberry, P., Orr-Ewing, D.C., O’Toole, L., Simpson, D. & Snell, N. 2003. Reintroduction and conservation of the red kite (Milvus milvus) in Britain: current threats and prospects for future range expansion. In Thompson, D.B.A., Redpath, S.M., Fielding, A.H., Marquiss, M. & Galbraith, C.A. (Eds.). Birds of Prey in a Changing Environment. The Stationary Office, Edinburgh. Pp. 407-416.

Clark, W., Duffy, K., Gorney, E., McGrady, M.J. & Schultz, C. 1986. Raptor ringing in Eilat, Israel.Sandgrouse 7: 21-28.

Cresswell, W., Holt, S., Reid, J.M., Whitfield, D.P. & Mellanby, R.J. 2003. Do energetic demands constrain incubation scheduling in a biparental species? Behavioral Ecology 14: 97-102.

Cresswell, W., Holt, S., Reid, J.M., Whitfield, D.P., Mellanby, R.J., Norton, D. & Waldron, S. 2004. The energetic costs of egg heating constrain incubation attendance but do not determine daily energy expenditure in the pectoral sandpiper. Behavioral Ecology 15: 498-507.

Cresswell, W., Lind, J., Quinn, J.L., Minderman, J. & Whitfield, D.P. 2007. Ringing or colour-banding does not increase predation risk in Redshank Tringa totanusJournal of Avian Biology 38, 309-316.

Cresswell, W. & Whitfield, D.P. 1994. The effects of raptor predation on wintering wader populations at the Tyninghame estuary, southeast Scotland. Ibis 136: 223-232.

Cresswell, W. & Whitfield, D.P. 2008. How starvation risk in redshanks Tringa totanus results in predation mortality from sparrowhawks Accipiter nisusIbis 150 (Suppl. 1): 209-218.

Downie, I.S., Foster, G.N., Coulson, J.C. & Whitfield, D.P. 1998. Distribution of aquatic macroinvertebrates within peatland pool complexes in the Flow Country, Scotland. Hydrobiologia 377: 95-105.

Downie, I.S., Foster, G.N., Coulson, J.C. & Whitfield, D.P. 1998. The water beetles of Badanloch Bog, Sutherland. Glasgow Naturalist 23: 31-34.

Duffy, K. 1988. Barn owls breeding near Edinburgh, 1985.  Lothian Bird Report 1987.  Scottish Ornithologists Club, Lothian Branch. P. 63-64.

Duffy, K. 1989. Some notes on night heron sightings in the Lothians. Lothians Bird Report 1988. Scottish Ornithologists Club. P. 75-76.

Duffy, K. 1990.  Breeding the hawk-headed parrot (Deroptyus acciptrinus acciptrinus) at Penscynor Wildlife Park. Association of British Wild Animal Keepers 17 (1): 14 –15.

Eaton, M.A., Dillon, I.A., Stirling-Aird, P.K. & Whitfield, D.P. 2007. Status of golden eagle Aquila chrysaetos in Britain in 2003. Bird Study 54: 212-220.

Evans, R., Broad, R., Duffy, K., MacLennan, A., Bainbridge, I. & Mudge, G. 2003. Re-establishment of a population of white-tailed eagles in Scotland. In Helander, B., Marquiss, M. & Bowerman, W. (Eds.).Sea Eagle 2000. Proceedings from an international conference at Bjorko, Sweden, 13 – 17 September 2000. Swedish Soc. for Nature Conservation/SNF, Stockholm. Pp. 397-403.

Evans, R.J., Douse, A., MacLennan, A.M., Sexton, D., Whitfield, D.P. & Wilson, J.D. 2009. Growth and demography of a re-introduced white-tailed eagle Haliaeetus albicilla population in Scotland. Ibis151: 244-254.

Evans, R. J., Pearce-Higgins, J., Whitfield, D. P., Grant, J. R., MacLennan, A. M. & Reid, R. 2010. Comparative nest site characteristics of sympatric White-tailed Haliaeetus albicilla and Golden EaglesAquila chrysaetos in western Scotland. Bird Study 57: 473-482.

Evans, S., Redpath, S.M., Leckie, F. & Mougeot, F. (2007) Alternative methods for estimating breeding density in an upland gamebird, the red grouse. Wildlife Biology. 13 (2): 130-139.

Fielding, A.H., Haworth, P.F., Morgan, D.H., Thompson, D.B.A. & Whitfield, D.P. 2003. The impact of golden eagles (Aquila chrysaetos) on a diverse bird of prey assemblage. In Thompson, D.B.A., Redpath, S.M., Fielding, A.H., Marquiss, M. & Galbraith, C.A. (Eds.). Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh. Pp. 221-244.

Fielding, A., McLeod, D.R.A., McGrady, M., Whitfield, D.P. & Haworth, P. 2003. Modelling the impact of land use change on golden eagles Aquila chrysaetos. In Thompson D. B. A., Redpath, S.M., Fielding, A.H., Marquiss, M. & Galbraith, C.A. (Eds.). Birds of Prey in a Changing Environment. The Stationary Office, Edinburgh. Pp. 323-240.

Fielding, A.H., Whitfield, D.P. &   McLeod, D.R.A.  2006. Spatial association as an indicator of the potential for future interactions between wind energy developments and golden eagles Aquila chrysaetos in Scotland. Biological Conservation 131: 359-369.

Fielding, A.H., Whitfield, D.P. & Whitehead, S. In press. Population dynamics of the hen harrier Circus cyaneus in Wales. Bird Study vol & pages 

Fuller, R.J., Gillings, S. & Whitfield, D.P. 1999. Responses of breeding birds to expansion of scrub in the eastern Scottish Highlands: preliminary implications for conservation strategies. Vogelwelt 120 Suppl.: 53-62.

Fuller, R. J. and Jackson, D. B. 1999. Changes in populations of breeding waders on the machair of North Uist, Scotland 1983 – 1998. Wader Study Group Bulletin 90: 47-55.

Galbraith, H., Murray, S., Duncan, K., Smith, R., Whitfield, D.P. & Thompson, D.B.A. 1993. Diet and habitat use of the dotterel Charadrius morinellus in Scotland. Ibis 135: 148-155.

Galbraith, H., Murray, S., Rae, S., Whitfield, D.P. & Thompson, D.B.A. 1993. Numbers and distribution of the dotterel (Charadrius morinellus) breeding in Great Britain. Bird Study 40: 161-169.


Haworth, P. F., McGrady, M.J., Whitfield, D. P., Fielding, A. H. & McLeod, D. R. A.  2006. Ranging distance of adult golden eagles Aquila chrysaetos in western Scotland according to season and breeding status. Bird Study 53: 265-273.


Holt, S. & Whitfield, D.P. 2003. Timing of nest departure by broods of dotterel Charadrius morinellus.Bird Study 49: 88-90.

Holt, S., Whitfield, D.P., Duncan, K., Rae, S. & Smith, R.D. 2002. Mass loss in incubating Eurasian dotterel: adaptation or constraint? Journal of Avian Biology 33: 219-224.

Holt, S., Whitfield, D.P. & Gordon, J. 2002. Potential reproductive rates in the Eurasian dotterel. Bird Study 49: 87-88.

Jackson, D.B., 1994. Breeding dispersal and site-fidelity in three monogamous wader species in the Western Isles, UK. Ibis 136: 463-473.

Jackson, D.B., 2001. Experimental removal of introduced hedgehogs improves wader nest success in the Western Isles, Scotland. Journal of Applied Ecology 38: 802-812.

Jackson D.B., 2003. Waders, hedgehogs and machair: research and conservation lessons from the Outer Hebrides. Wader Study Group Bulletin Vol. 100: 14-19.

Jackson D.B., 2003. Between-lake differences in the diet and provisioning behaviour of black-throated divers Gavia arctica breeding in Scotland. Ibis 145: 30-44.

Jackson, D.B., 2005. Environmental correlates of lake occupancy and brood survival of black-throated divers Gavia arctica in Scotland. Bird Study 52: 225-236.

Jackson, D.B., 2006. The breeding biology of introduced hedgehogs on a Scottish island: lessons for population control and bird conservation. Journal of Zoology (London) 268: 303-314.

Jackson , D.B. 2006. Factors affecting the abundance of introduced hedgehogs (Erinaceus europaeus) to the Hebridean island of South Uist in the absence of natural predators and the implications for nesting birds. Journal of Zoology (London) 271(2): 210-217. 

Jackson, D.B., Fuller, R.J. and Campbell, S.T. 2004 Long-term population changes among breeding shorebirds in the Outer Hebrides, Scotland, in relation to an introduced mammalian predator. Biological Conservation 117: 151-166.

Jackson, D.B. & Green, R. E. 2000. The importance of the introduced hedgehog (Erinaceus europaeus) as a predator of the eggs of waders (Charadrii) on the machair in South Uist, Scotland.Biological Conservation 93: 33-348.

Janss, G. F. E, de Lucas, M., Lazo, A., Ferrer, M. & Whitfield, D. P. 2010. The precautionary principle and wind-farm planning in Andalucía. Biological Conservation 143: 1827-1828.

Jones, C.G. & Duffy, K. 1991. Field work on the echo parakeet. Psittascene 3 (4): 6-7.

Jones, C.G. & Duffy, K. 1992. Conservation of the echo parakeet Psittacula eques from Mauritius.Psittascene 4 (4): 7-10.

Jones, C.G. & Duffy, K. 1993. Conservation of the echo parakeet Psittacula eques. J. Wildlife Preservation Trusts 29: 126-148.

Leckie, F., Arroyo, B.E., Thirgood. S.J. & Redpath, S.M. (2008) Parental differences in brood provisioning by hen harriers Circus cyaneusBird Study 55: 209-215.

Leckie, F. and Campbell, S. (2000) A new record of successful tree nesting peregrines. Scottish Birds21(1): 45-46.

Leckie, F. M., Thirgood, S.J., May, R., & Redpath S. M. (1998) Variation in the diet of red foxes on Scottish moorland in relation to prey abundance. Ecography 21(6): 599-604.

de Lucas, M., Janss, G.F.E., Whitfield, D.P. & Ferrer, M. 2008. Collision fatality of raptors in wind farms does not depend on raptor abundance. Journal of Applied Ecology 45: 1695-1704.

Madders, M., Leckie, F.M., Watson, J., & McKay, C.R. (1998) Distribution and foraging habitat preferences of choughs on The Oa peninsula, Islay. Scottish Birds 19 (5): 280-289.

Madders, M. & Whitfield, D.P. 2006. Upland raptors and the assessment of windfarm impacts. Ibis 148 Supplement: 43-56.


McLeod, D. R. A., Whitfield, D. P., Fielding, A. H., Haworth, P. & McGrady M.J. 2003. Predicting home range use by golden eagles Aquila chrysaetos in western Scotland. Avian Science 2: 183-198.


McLeod, D. R. A., Whitfield, D.P. & M.J. McGrady 2002. Improving prediction of golden eagle (Aquila chrysaetos) ranging in western Scotland using GIS and terrain modelling. Journal of Raptor Research36 (1 Suppl.), March 2002: 70-77.

Mee, A., Whitfield, D.P., Thompson, D.B.A. & Burke, T.A. 2004. Extra-pair paternity in the common sandpiper Actitis hypoleucos, revealed by DNA fingerprinting. Animal Behaviour 67: 333-342.

Moss, R., Biggins, A., Leckie, F., Poole, T., Gordon, J., Leslie, C. & Kortland, K. May 2010. The distribution of capercaillie droppings in relation to sources of disturbance: part 3 Glenmore Forest Park. Progress Report to Capercaillie BAP group, draft.

Moss, R., Biggins, A., Leckie, F., Jones, G., Jones, S. & Lambie, D. October 2010. The distribution of capercaillie droppings in relation to sources of disturbance: part 2 Boat of Garten Wood. Progress Report to Capercaillie BAP group, draft. Version 2.

Mougeot, F., Dawson, A., Redpath, S.M. & Leckie, F. (2005) Testosterone and autumn territorial behaviour in male red grouse Lagopus lagopus scoticusHormones & Behavior 47: 576-584.

Mougeot, F., Piertney, S., Leckie, F., Evans, S., Moss, R., Redpath, S.M. & Hudson, P.J. (2005) Experimentally increased aggressiveness reduces population kin structure and subsequent recruitment in red grouse Lagopus lagopus scoticus. Journal of Animal Ecology 74: 488-497.

Mougeot, F., Redpath, S.M., Leckie, F. and Hudson, P.J. (2003) The effect of aggressiveness on the population dynamics of a territorial bird. Nature 421: 737-739.

Mougeot, F., Redpath, S.M. & Leckie, F. (2005) Ultra-violet reflectance of male and female red grouse sexual ornaments reflects nematode parasite intensity. Journal of Avian Biology 36: 203-209.

O’Brien, M., Tharme, A. & Jackson, D. 2002. Changes in breeding wader numbers on Scottish farmed land during the 1990s. Scottish Birds 23: 10-21.

Parrott, D., Henderson, I., Deppe, C. & Whitfield, P. 2008. Scottish racing pigeons killed by peregrine falcons Falco peregrinus: estimation of numbers from ring recoveries and peregrine daily food intake.Bird Study 55: 32-42.



Piertney, S.B., Lambin, X., McColl, A., Lock, K., Bacon, P., Dallas, J., Mougeot, F., Leckie, F., Racey, P., Redpath, S.M. & Moss, R. 2008. Temporal changes in kin structure through a population cycle in a territorial bird, the red grouse (Lagopus lagopus scoticus). Molecular Ecology  17: 2544-2551.




Redpath, S. Amar, A., Madders, M., Leckie, F. & Thirgood, S. (2002) Hen harrier foraging success in relation to land use in Scotland. Animal Conservation 5, 113-118


Redpath, S.M., Arroyo, B.E., Etheridge, B., Leckie, F., Bouwman, K. & Thirgood, S.J. (2002) Temperature and hen harrier productivity: from local mechanisms to geographical patterns. Ecography25: 533-540.

Redpath, S.M., Arroyo, B.E., Leckie, F.M., Bacon, P., Bayfield, N., Gutiérrez, R.J. & Thirgood, S.J. (2004) Using decision modelling with stakeholders to reduce human-wildlife conflict: a raptor - grouse case study. Conservation Biology 18: 350-359.

Redpath, S.M., Leckie, F., Arroyo, B.., Amar, A. & Thirgood, S. (2006) Compensating for the costs of polygyny in hen harriers Circus cyaneusBehavioural Ecology and Sociology 60 (3): 386-391.

Redpath, S.M., Mougeot, F., Leckie, F.M., Elston, D.A., & Hudson, P.J. (2006) Testing the role of parasites in driving the cyclic populatin of a gamebird. Ecology Letters 9: 410-418.

Redpath, S., Mougeo,t F., Leckie, F. & Evans, S. (2006) The effects of autumn testosterone on survival and productivity in red grouse Lagopus lagopus scoticusAnimal Behaviour 71:1297-1305.

Redpath, S.M., Thirgood, S.J. & Leckie, F. (2001) Does supplementary feeding reduce predation of red grouse by hen harriers? J. Appl. Ecol. 38 (6): 1157-1168.

Reid, J.M., Cresswell, W., Holt, S., Mellanby, R.J., Whitfield, D.P. & Ruxton, G.D. 2002. Nest scrape design and clutch heat loss in pectoral sandpipers Calidris melanotosFunctional Ecology 16: 305-312.

Scott, D. 1998. Impact of red deer on a Scots pine plantation after the removal of deer fencing.Scottish Forestry 52: 8-13.

Scott, D., Welch, D. Thurlow, M., Elston, D.A. 2000. Regeneration of Pinus sylvestris in a natural pinewood in N.E. Scotland following reduction in grazing by Cervus elaphus. Forest Ecology and Management 130: 199-211.

Scott, D. 2002. Recovery of a dwarf birch (Betula nana) population following reduction in grazing by red deer (Cervus elaphus). Botanical Journal of Scotland 53(2): 155-167.

Scott, D., Bayfield, N.G., Cernusca, A., Elston, D.A. 2002. Use of a weighing lysimeter system to assess the effects of trampling on evapo-transpiration of montane plant communities. Canadian Journal of Botany 80: 678-683.

Scott, D., Welch, D., Van der Wal, R., Elston, D.A. 2007. Response of the montane moss Racomitrium lanuginosum to changes in sheep grazing and snow-lie. Applied Vegetation Science 10: 229-238.

Scott, D., Welch, D., Elston, D.A. 2009. Long-term effects of leader browsing by deer on the growth of Sitka spruce (Picea sitchensis). Forestry 82: 387-401.

Seivwright, L., Redpath, S.M., Mougeot, F., Leckie, F. & Hudson, P.J. 2005. Interactions between intrinsic and extrinsic mechanisms in a cyclic species: testosterone increases parasite infection in red grouse. Proceedings of the Royal Society-Biological Sciences 172: 2299-2304.

Smart, J., Amar, A., Sim, I., Etheridge, B., Cameron, D., Christie, G. & Wilson, J. 2010.  Illegal killing slows population recovery of Red Kite.  Biological Conservation 143: 1278-1286.

Terraube, J., Arroyo, B., Mougeot, F., Madders, M. & Bragin, E. 2009. Interacciones entre estrategias de reproducción y trofica en dos especies de rapaces simpátricas: los aguiluchos papialbo (Circus macrourus) y cenizo (Circus pygargus) en el norte de Kazajstán. In “Conservación y situación poblacional de los aguiluchos en Eurasia”. (Alarcos S. &  Palacios, M.J. eds.). Junta de Extremadura.

Terraube, J., Arroyo, B.E., Mougeot, F., Madders, M., Watson, J. & Bragin, E. 2009. Breeding biology of the pallid harrier Circus macrourus in north-central Kazakhstan: implications for the conservation of a Near-Threatened species. Oryx 43 (1): 104-112. 



Thirgood, S.J., Leckie, F.M. & Redpath, S.M.  (1996) Diurnal and seasonal variation in line transect counts of moorland passerines. Bird Study 42: 257-259.


Thompson, D.B.A., Whitfield, D.P., Galbraith, H., Duncan, K., Smith, R.D., Murray, S. & Holt, S. 2003. Breeding bird assemblages of alpine areas in Scotland. In Nagy, L., Grubherr, G., Körner, Ch. & Thompson, D.B.A. (Eds.) Alpine Biodiversity in Europe, Springer-Verlag, Berlin. Pp. 327-338.

Van der Wal, R., Pearce, I.S.K., Brooker, R., Scott, D., Welch, D., Woodin, S. 2003. Interplay between nitrogen deposition and grazing accelerates habitat degradation. Ecology Letters 6: 141-146.

Walker D.G. 1983. Golden eagle killing mobbing carrion crows. Brit. Birds 76: 309-310.

Walker, D.G. 1987 Observations on the post-fledging period of the golden eagle Aquila chrysaetos in England. Ibis 129: 92-96.

Walker D.G. 1988 Unusual golden eagle call. Scott. Bird News 9: 8.

Walker, D.G. 1988 The behaviour and movements of a juvenile golden eagle Aquila chrysaetos in England. Ibis 130: 564-565.

Walker, D.G. 2000. Food supply and breeding performance of the Haweswater golden eagles. Birds and Wildlife in Cumbria: 65-68.

Walker, D.G. 2003. Some notes on the red deer (Cervus elaphus) at Haweswater, Mardale. The Carlisle Naturalist 11: 14-16.

Walker, D.G. 2004 The Lakeland Eagles. ISBN 0 9517378 0 5. David Walker.

Walker, D.G. 2009. Call of the Eagle. Whittles Publishing, UK.

Walker, D., McGrady, M., McCluskie, A., Madders, M. & McLeod, D.R.A. 2005. Resident golden eagle ranging behaviour before and after construction of a windfarm in Argyll. Scottish Birds 25 (1): 24-40.

Watson, J., Fielding, A.H., Whitfield, D.P., Broad, R.A., Haworth, P., Nellist, K., Crane, K. & Macdonald, E.J. 2003. Golden eagle breeding performance in relation to climate in western Scotland during the period 1981-2000. In Thompson, D.B.A., Redpath, S.M., Fielding, A.H., Marquiss, M. & Galbraith, C.A. (Eds.) Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh. Pp. 149-161.

Watson, J. & Whitfield, P. 2002. A conservation framework for the golden eagle Aquila chrysaetos in Scotland. Journal of Raptor Research 36 (1 Supplement): 41-49.

Welch, D., Scott, D. 1985. Further observations on a Callunetum intensively grazed by mountain hares (Lepus timidus L.). Trans. Bot. Soc. Edinb. 44: 323-333.

Welch. D., Staines. B.W., Scott. D., Catt. D.C. 1987. Bark stripping damage by red deer in a Sitka spruce forest in Western Scotland. I. Incidence. Forestry 60: 249-262.

Welch. D., Chambers. M.G., Scott. D., Staines. B.W. 1988. Roe-deer browsing on spring-flush growth of Sitka spruce. Scottish Forestry 42: 33-43.

Welch. D., Staines. B.W., Scott. D., Catt. D.C. 1988. Bark stripping damage by red deer in a Sitka spruce forest in Western Scotland. II. Wound size and position. Forestry 61: 245-254.

Welch. D., Staines. B.W., Catt. D.C., Scott. D. 1990. Habitat usage by red (Cervus elaphus) and roe (Capreolus capreolus) deer in a Scottish Sitka spruce plantation. Journal of Zoology London: 221, 453-476.

Welch. D., Staines. B.W., Scott. D., Catt. D.C., French. D.D. 1991. Leader browsing by red and roe deer on young Sitka spruce trees in Western Scotland I. Damage rates and the influence of habitat factors. Forestry 64: 61-82.

Welch. D., Staines. B.W., Scott. D., French. D.D. 1992. Leader browsing by red and roe deer on young Sitka spruce trees in Western Scotland. II. Effects on growth and tree form. Forestry 65: 309-330.

Welch. D., Scott. D., Staines B.W. 1995. Survival rates and performance of multi-trunked trees in even-aged stands of Sitka spruce in Western Scotland. Forestry 68: 245-253.

Welch. D., Scott. D. 1995. Studies in the grazing of heather moorland in North-East Scotland VI. 20-year trends. Journal of Applied Ecology 32: 596-611.

Welch, D., Scott, D., Staines, B.W. 1997. Bark stripping damage by red deer in a Sitka spruce forest in Western Scotland. III. Trends in wound condition. Forestry 70: 113-120.

Welch. D., Scott, D. 1997. Decline of moorland plants following the establishment of a Scots pine plantation. Botanical Journal of Scotland 49: 27-37.

Welch, D., Scott, D., Staines, B.W. 1998. Bark stripping damage by red deer in a Sitka spruce forest in Western Scotland. IV. Survival and performance of wounded trees. Forestry 71: 225-235.

Welch, D., Scott, D., Doyle, S. 2000. Studies in the paradox of seedling rarity in Vaccinium myrtillus L. in North-East Scotland. Botanical Journal of Scotland 52: 17-30.

Welch, D., Scott, D. 2001. Seasonal and spatial patterns in sheep grazing on a high-level plateau in the Grampian mountains, Scotland. Scottish Geographical Journal 116: 299-314.

Welch, D. Scott, D., Thompson, D.B.A. 2005. Changes in the composition of Carex bigelowii-Racomitrium lanuginosum moss heath on Glas Maol, Scotland, in response to sheep grazing and snow fencing. Biological Conservation 122: 621-631.

Welch, D. Scott, D, Mitchell, R., Elston  D. 2007. Differential recovery of heather Calluna vulgaris in   Scottish moorland after easing of heavy grazing pressure from red deer Cervus elaphusBotanical Journal of Scotland 58: 1-17.

Welch, D., Scott, D. 2008. An estimate of timber degrade due to bark-stripping in Sitka spruce in a Scottish plantation. Forestry 81: 489-497.

Whitfield, D.P. 1985. Raptor predation on wintering waders in southeast Scotland. Ibis 127: 544-558.

Whitfield, D.P. 1986. Plumage variability and territoriality in breeding turnstone Arenaria interpres: status signalling or individual recognition? Animal Behaviour 34: 1471-1482.

Whitfield, D.P. 1987. Plumage variability, status signalling and individual recognition in avian flocks.Trends in Ecology and Evolution 2: 13-17.

Whitfield, D.P. 1990. Male choice and sperm competition as constraints on polyandry in the red-necked phalarope Phalaropus lobatusBehavioural Ecology & Sociobiology 27: 247-254.

Whitfield, D.P. 1990. Individual feeding specialisations in wintering turnstone Arenaria interpres.Journal of Animal Ecology 59: 193-211.

Whitfield, D.P. 1995. Behaviour and ecology of a polyandrous population of grey phalaropesPhalaropus fulicarius in Iceland. Journal of Avian Biology 26: 349-352.

Whitfield, D.P. 1997. Waders (Charadrii) on Scotland’s blanket bogs: recent changes in numbers of breeding birds. In Parkyn, L., Stoneman, R.E. & Ingram, H.A.P. (Eds.). Conserving Peatlands. CAP International, Wallingford. Pp. 103-111.

Whitfield, D.P. 1988. Sparrowhawks Accipiter nisus affect the spacing behaviour of wintering turnstone Arenaria interpres and redshank Tringa totanusIbis 130: 284-287.

Whitfield, D.P. 1988. The social significance of plumage variability in wintering turnstone Arenaria interpresAnimal Behaviour 36: 408-415.

Whitfield, D.P. 1999. Methods for ageing dotterel Charadrius morinellusRinging & Migration 19: 200-204.

Whitfield, D.P. 2002. Status of breeding dotterel Charadrius morinellus in Britain in 1999. Bird Study49: 237-249.

Whitfield, D.P. 2003. Does timing of breeding affect breeding plumage acquisition in shorebirds?Wader Study Group Bulletin 101/102: 77-80.

Whitfield, D.P. 2003. Raptor predation on non-breeding shorebirds: some thoughts for the future.Wader Study Group Bulletin 100: 134-137.

Whitfield, D.P. 2003. Density-dependent mortality of wintering dunlins Calidris alpina through predation by Eurasian sparrowhawks Accipiter nisusIbis 145: 432-438.

Whitfield, D.P. 2003. Predation by Eurasian sparrowhawks produces density-dependent mortality in wintering redshanks. Journal of Animal Ecology 72: 27-35.

Whitfield, D.P. 2003. Redshank Tringa totanus flocking behaviour, distance from cover and vulnerability to sparrowhawk Accipiter nisus predation. Journal of Avian Biology 34: 163-169.

Whitfield, D.P. 2003. The effects of predation by sparrowhawks on the behaviour and population dynamics of wintering redshanks. In Thompson, D.B.A., Redpath, S.M., Fielding, A.H., Marquiss, M. & Galbraith, C.A. (Eds.). Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh. Pp. 269-274.

Whitfield, D.P. & Brade, J.J. 1991. The breeding behaviour of the knot Calidris canutusIbis 133: 246-255.

Whitfield, D.P., Brade, J.J., Burton, R.W., Hankinson, K.W. & Young, S. 1996. The abundance of breeding knot Calidris canutus islandicaBird Study 43: 290-299.

Whitfield, D.P. & Coupar, A. 2008. Effects of terrestrial wind farms on birds and upland habitats. InGalbraith, C.A. & Baxter, J.M. (Eds). Energy and the Natural Heritage. The Stationery Office, Edinburgh. Pp. 205-220.

Whitfield, D.P., Cresswell, W., Ashmole, N.P., Clark, N.A. & Evans, A.D. 1999. No evidence for sparrowhawks selecting redshanks according to size or condition. Journal of Avian Biology 30: 31-39.

Whitfield, D.P., Evans, R.J., Broad, R.A., Fielding, A.H., Haworth, P.F., Madders, M. & McLeod, D.R.A.2002. Are reintroduced white-tailed eagles in competition with golden eagles? Scottish Birds 23: 36-45.

Whitfield, D.P., Douse, A., Evans, R.J., Grant, J., Love, J., McLeod, D.R.A., Reid, R. & Wilson, J.D. 2009. Natal and breeding dispersal in a reintroduced population of white-tailed eagles Haliaeetus albicillaBird Study 56: 177-187.

Whitfield, D.P., Duffy, K., McLeod, D.R.A., Evans, R.J., MacLennan, A.M., Reid, R., Sexton, D., Wilson, J.D. & Douse, A. 2009. Juvenile dispersal of white-tailed eagles in western Scotland. Journal of Raptor Research 43: 110-120.

Whitfield, D.P., Duncan, K., Pullan, D. & Smith, R.D. 1996. Recoveries of Scottish-ringed dotterelCharadrius morinellus in the non-breeding season: evidence for seasonal shifts in wintering distribution: Ringing & Migration 17: 105-110.

Whitfield, D.P., Evans, A.D. & Whitfield, P.A. 1988. The impact of raptor predation on wintering waders. Proceedings of the International Ornithological Congress 19: 674-687.

Whitfield, D.P., Fielding, A.H., Gregory, M.J.P., Gordon, A.G., McLeod, D.R.A. & Haworth, P.F. 2007. Complex effects of habitat loss on golden eagles Aquila chrysaetosIbis 149: 26-36.

Whitfield, D.P., Fielding, A.H., McLeod, D.R.A. & Haworth, P.F. 2004. Modelling the effects of persecution on the population dynamics of golden eagles in Scotland. Biological Conservation 119: 319-333.

Whitfield, D.P., Fielding, A.H., McLeod, D.R.A. & Haworth, P.F. 2004. The effects of persecution on age of breeding and territory occupation in golden eagles in Scotland. Biological Conservation 118: 249-259.

Whitfield, D.P., Fielding, A.H., McLeod, D.R.A., Haworth, P.F. & Watson, J. 2006. A conservation framework for the golden eagle in Scotland: refining condition targets and assessment of constraint influences. Biological Conservation 130: 465-480.

Whitfield, D.P., Fielding, A.H., McLeod, D.R.A., Morton, K., Stirling-Aird, P. & Eaton, M. 2007. Factors constraining the distribution of golden eagles Aquila chrysaetos in Scotland. Bird Study 54: 199-211.

Whitfield, D.P., Fielding, A.H. & Whitehead, S. 2008. Long-term increase in the fecundity of hen harriers Circus cyaneus in Wales is explained by reduced human interference and warmer weather.Animal Conservation 11: 144-152.

Whitfield, D.P. & Magnusson, J. 1987. The migration of high arctic waders through Melrakkasletta, northeast Iceland. Wader Study Group Bulletin, 49 Suppl./IWRB Special Publication 7: 85-89.

Whitfield, D.P., McLeod, D.R.A., Fielding, A.H., Evans, R.J., Broad, R.A. & Haworth, P.F. 2001. The effects of forestry on golden eagles on the island of Mull, western Scotland. Journal of Applied Ecology38: 1208-1220.

Whitfield, D.P., McLeod, D.R.A., Fielding, A.H., Watson, J. & Haworth, P.F. 2003. The association of grouse moors in Scotland with the illegal use of poisons to control predators. Biological Conservation114: 157-163.

Whitfield, D.P., Reid, R., Haworth, P.F., Madders, M., Marquiss, M., Tingay, R.E. & Fielding, A.H. 2009. Diet specificity is not associated with increased reproductive performance of golden eagles Aquila chrysaetos in western Scotland. Ibis 151: 255-264.

Whitfield, D.P., Ruddock, M. & Bullman, R. 2008. Expert opinion as a tool for quantifying bird tolerance to human disturbance. Biological Conservation 141: 2708-2717.

Whitfield, D.P. & Tomkovich, P.S. 1996. Mating system and timing of breeding in Holarctic waders.Biological Journal of the Linnaean Society 57: 277-290.

Wotton, S.R., Carter, I., Cross, I.V., Etheridge, B.E. Snell, N., Duffy, K., Thorpe, R. & Gregory, R.D. 2002. Breeding status of the red kite Milvus milvus in Britain in 2000. Bird Study 49: 278-286.

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